1. School of Animal Science Yangtze University Jingzhou China
2. Research Center of Fishery Resources and Environment Guizhou University Guiyang China
3. Guangdong Provincial Key Laboratory of Marine Biotechnology Shantou University Shantou China
4. Key Lab of Freshwater Biodiversity Conservation Ministry of Agriculture Yangtze River Fisheries Research InstituteChinese Academy of Fishery Sciences Wuhan China