1. Department of Biological Sciences; Macquarie University; Sydney NSW Australia
2. Pied Babbler Research Project, Kuruman River Reserve; Van Zylsrus South Africa
3. Institute of Evolutionary Biology; School of Biological Sciences; University of Edinburgh; Edinburgh UK
4. DST/NRF Centre of Excellence at the Percy FitzPatrick Institute; University of Cape Town; Cape Town Western Cape South Africa
5. Centre for Evolutionary Biology; School of Animal Biology; University of Western Australia; Perth WA Australia