1. Department of Animal Behaviour Bielefeld University Konsequenz 45 Bielefeld 33615 Germany
2. Evolutionary Biology Bielefeld University Konsequenz 45 Bielefeld 33615 Germany
3. Conservation Ecology Group, Groningen Institute for Evolutionary Life Sciences (GELIFES) University of Groningen Nijenborgh 7 Groningen 9747AG The Netherlands
4. Centre for Ecology and Conservation University of Exeter (Penryn Campus) Penryn Cornwall TR10 9FE United Kingdom