1. The Sainsbury Laboratory Norwich Research Park Norwich NR4 7UH UK
2. Department of Plant–Microbe Interactions Max Planck Institute for Plant Breeding Research Carl‐von‐Linné weg 10 50829 Köln Germany
3. Wageningen UR Plant Breeding Wageningen University and Research Centre Droevendaalsesteeg 1 Wageningen 6708 PB the Netherlands
4. Key Laboratory of Horticultural Plant Biology College of Life Science and Technology, Ministry of Education National Center for Vegetable Improvement (Central China) Potato Engineering and Technology Research Center of Hubei Province Huazhong Agricultural University Wuhan 430070 Hubei China