1. INRA; UMR 1202 BIOGECO; F-33610 Cestas France
2. Univ. Bordeaux; BIOGECO; UMR 1202; F-33400 Talence France
3. School of Biological Sciences; Royal Holloway University of London; Egham Surrey TW20 0EX UK
4. Scion (New Zealand Forest Research Institute); Christchurch 8540 New Zealand