1. School of Biological Sciences; University of Queensland; Brisbane Qld 4072 Australia
2. Department of Biology; Paleoecological Environmental Assessment and Research Lab (PEARL); Queen's University; Kingston ON K7L 3N6 Canada
3. School of Life Sciences; Silwood Park Imperial College London; Ascot Berkshire SL5 7PY UK
4. Zoology Department; School of Natural Sciences; Trinity College Dublin; Dublin 2 Ireland