1. Department of Biological Sciences; The University of Waikato; Private Bag 3105; Hamilton; 3240; New Zealand
2. Department of Ecology, Lincoln University; Burns Building Rm 520; Lincoln 7647; Christchurch; New Zealand
3. Institute of Biology Leiden; Leiden University; Sylviusweg 72 2333BE; PO Box 9505, 2300RA; Leiden; The Netherlands