1. Section for Biodiversity & Conservation; Department of Bioscience; Aarhus University; Grenåvej 14 8410 Kalø Denmark
2. School of Geography, Earth and Environmental Sciences; University of Birmingham; Birmingham UK
3. Section for Wildlife Ecology; Department of Bioscience; Aarhus University; Kalø Denmark
4. Section for Nature Conservation; The Nature Agency, Ministry of Environment and Food; Copenhagen Denmark
5. UR “Ecologie et dynamique des systèmes anthropisés” (EDYSAN, FRE 3498 CNRS-UPJV); Université de Picardie Jules Verne; Amiens France