1. Laboratoire d'Ecologie Alpine; CNRS; Université Grenoble Alpes; 2233 rue de la Piscine 38041 Grenoble France
2. UMR 7204 MNHN-UPMC-CNRS Conservation des Espèces; Restauration et Suivi des Populations; 55 rue Buffon 75005 Paris France
3. Research Institute for Nature and Forest; Kliniekstraat 25 1070 Brussel Belgium
4. Department of Geography; College of Life and Environmental Sciences; University of Exeter; Rennes Drive EX4 4RJ Exeter UK
5. Irstea; National Research Institute of Science and Technology for Environment and Agriculture; 2 rue de la Papeterie 38402 Saint Martin d'Hères France
6. Sustainability and Climate Change Programme; European Forest Institute; Yliopistokatu 6 80100 Joensuu Finland