1. Evolutionary Ecology Group; Department of Biology; University of Antwerp, Groenenborgerlaan 171, 2020; Antwerp Belgium
2. Institute of Biological and Environmental Sciences; University of Aberdeen; Tillydrone Avenue Aberdeen, AB24 2TZ UK
3. Terrestrial Ecology Unit; Department of Biology; Ghent University; K.L. Ledeganckstraat 35, 9000 Ghent Belgium