1. Department of Life Sciences; Imperial College London; Buckhurst Rd Ascot SL5 7PY UK
2. Centre for Environment, Fisheries and Aquaculture Science; Lowestoft Suffolk NR33 0HT UK
3. School of Environmental Sciences; University of East Anglia; Norwich Research Park Norwich NR4 7TJ UK
4. Department of Ecology and Evolutionary Biology and Kansas Biological Survey; University of Kansas; Lawrence KS 66047 USA
5. Laboratory of Populations; Rockefeller University; 1230 York Avenue New York NY 10065 USA