1. School of BioSciences; The University of Melbourne; Parkville Vic. 3010 Australia
2. Division of Biology; Imperial College London; Silwood Park Ascot SL5 7PY UK
3. Institute of Zoology; Zoological Society of London; Regent's Park London NW1 4RY UK
4. Centre for Biodiversity and Environment Research; University College London; Gower Street London WC1 E6BT UK
5. School of Life and Environmental Sciences; Deakin University; Burwood Vic. 3125 Australia