1. Marine Biology and Ecology Research Centre; Plymouth University; Drake Circus; Plymouth; PL4 8AA; UK
2. Institut Mediterrani d'Estudis Avançats IMEDEA (CSIC-UIB); Miquel Marqués 21; 07190; Esporles; Mallorca; Spain
3. Netherlands Institute for Sea Research (NIOZ); PO Box 59; 1790 AB; Den Burg; Texel; The Netherlands