1. Department of Conservation Biology; Doñana Biological Station (CSIC); C/Américo Vespucio s/n E-41092 Seville Spain
2. Department of Biology of Organisms and Systems; University of Oviedo; C/Valentín Andrés Álvarez s/n E-33071 Oviedo Spain
3. Unité Mixte de Recherche CNRS 5558 - Laboratoire de Biométrie et Biologie Evolutive; Université Claude-Bernard Lyon 1; 43 Boulevard du 11 novembre 1918 69622 Villeurbanne Cedex France
4. Department of Zoology & Entomology; University of the Free State; Private Bag X13 9866 Phuthaditjhaba South Africa