1. Department of Animal Ecology; Netherlands Institute of Ecology (NIOO-KNAW); PO Box 50 Wageningen 6700 AA the Netherlands
2. Computational Geo-Ecology; Institute for Biodiversity and Ecosystem Dynamics; University of Amsterdam; Science Park 904 Amsterdam 1098 XH the Netherlands
3. Department of Migration and Immuno-Ecology; Max Planck Institute for Ornithology; Am Obstberg 1 Radolfzell D-78315 Germany
4. Department of Biology; University of Konstanz; Konstanz D-78457 Germany
5. VHL University of Applied Sciences; PO Box 9001 Velp 6880 GB the Netherlands