1. Aquatic Ecology and Environmental Biology; Institute for Water and Wetland Research; Radboud University Nijmegen; NL-6525; AJ Nijmegen; The Netherlands
2. Deltares Subsurface and Groundwater Systems; P.O. Box 85467; NL-3584; CB Utrecht; The Netherlands
3. Institute of Biodiversity and Ecosystem Dynamics; University of Amsterdam; P.O. Box 94248; NL-1090; GE Amsterdam; The Netherlands