1. CAS Key Laboratory of Tropical Marine Bio-resources and Ecology(LMB), Guangdong Provincial Key Laboratory of Applied Marine Biology (LAMB), South China Sea Institute of Oceanology; Chinese Academy of Sciences; 164 West Xingang Road Guangzhou 510301 China
2. University of Chinese Academy of Sciences; Beijing 100049 China
3. South China Sea Bio-Resource Exploitation and Utilization Collaborative Innovation Center; Guangzhou China