1. Yangtze River Fisheries Research Institute; Chinese Academy of Fishery Sciences; Wuhan 430223 China
2. Freshwater Fisheries Research Center; Chinese Academy of Fishery Sciences; Wuxi 214081 China
3. Hu Bei Freshwater Aquaculture Collaborative Innovation Center; Wuhan 430070 China
4. School of Chemical Engineering & Biotechnology; XingTai University; Xingtai 054001 China