1. Department of Pathology; Shiga University of Medical Science; Otsu Shiga 520-2192 Japan
2. Joint Department of Veterinary Medicine, Faculty of Agriculture; Tottori University; Tottori 680-8553 Japan
3. Laboratory of Microbiology, Graduate School of Veterinary Medicine; Hokkaido University; Sapporo 060-0818 Japan
4. Global Station for Zoonosis Control, Global Institution for Collaborative Research and Education (GI-CoRE); Hokkaido University; Sapporo Hokkaido 060-0815 Japan
5. School of Veterinary Medicine; Kitasato University; Towada Aomori 034-8628 Japan
6. Research Center for Animal Life Science; Shiga University of Medical Science; Otsu Shiga 520-2192 Japan
7. Research Center for Zoonosis Control; Hokkaido University; Sapporo Hokkaido 001-0020 Japan