1. Centre for Sustainable Ecosystem Solutions; School of Biological Sciences; University of Wollongong; Wollongong NSW 2522 Australia
2. Centre for Evolutionary Biology; University of Western Australia; Crawley WA 6008 Australia
3. School of Science and Health; Western Sydney University; Hawkesbury Richmond NSW 2751 Australia
4. Department of Ecology, Evolution, and Behavior; University of Minnesota; Saint Paul MN 55108 USA