1. Department of World Heritage; Cyber University; Nagoya Office, Ikegami-cho 2-7-1, Ikegami Jyutaku R203, Chikusa-ku Nagoya 464-0029 Japan
2. Institute of Biodiversity Science; Fudan University; Handan Road 220 Shanghai 200433 China
3. Department of Life Sciences; Natural History Museum; London SW7 5BD UK