1. School of Life and Environmental Sciences; Faculty of Science; The University of Sydney; RMC Gunn Building (B19) NSW 2006 Australia
2. San Diego Zoo Global; PO Box 120551 San Diego CA 92112 USA
3. Department of Biology; University of Hawai'i at Hilo; 200 West Kāwili Street Hilo HI 96720 USA
4. School of Biological Sciences; University of Canterbury; Private Bag 4800 Christchurch New Zealand
5. Allan Wilson Centre for Molecular Ecology and Evolution; Department of Zoology; University of Otago; PO Box 56 Dunedin 9054 New Zealand