1. Evolutionary Genomics Group; Department of Botany and Zoology; Stellenbosch University; Private Bag X1 Stellenbosch 7602 South Africa
2. Department of Ichthyology and Fisheries Sciences; Rhodes University; P.O. Box 94 Grahamstown 6140 South Africa
3. National Marine Information and Research Centre; Private Bag 912 Swakopmund Namibia
4. Molecular Ecology and Evolution Programme; Department of Genetics; University of Pretoria; Private Bag X20 Hatfield, Pretoria 0028 South Africa