1. Department of Forest Science Graduate School of Agriculture, Hokkaido University Sapporo Hokkaido Japan
2. Tomakomai Research Station Field Science Center for Northern Biosphere, Hokkaido University Tomakomai Japan
3. Center for Environmental Biology and Ecosystem Studies National Institute for Environmental Studies Ibaraki Japan
4. Graduate School of Environmental Science Hokkaido University Sapporo Hokkaido Japan
5. Department of Biological Sciences School of Science, Hokkaido University Sapporo Hokkaido Japan