1. Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany; Chinese Academy of Sciences; Kunming Yunnan 650201 China
2. Woodrow Wilson School of Public and International Affairs; Princeton University; Princeton NJ 08544 USA
3. State Key Laboratory of BioControl, College of Ecology and Evolution/School of Life Sciences; Sun Yat-Sen University; Guangzhou Guangdong 510275 China
4. World Agroforestry Center; East and Central Asia; Kunming Yunnan 650201 China
5. Department of Ecology and Evolutionary Biology; Princeton University; Princeton NJ 08544 USA