1. Faculty of Agriculture; Tokyo University of Agriculture and Technology; Fuchu Tokyo 183-8509 Japan
2. School of Biological Sciences; The University of Queensland; Brisbane Qld 4072 Australia
3. Center for Environmental Biology and Ecosystem Studies; National Institute for Environmental Studies; Tsukuba Ibaraki 305-8506 Japan
4. Department of Integrative Biology; University of Guelph; Guelph Ontario N1G 2W1 Canada
5. The Nature Conservation Society of Japan; 1-16-10 Shinkawa Chuo-ku Tokyo 104-0033 Japan