1. School of Biological Sciences; Monash University; VIC 3800 Australia
2. Organisation for Tropical Studies; Scientific Services; Kruger National Park Mpumalanga 1350 South Africa
3. Global Change and Sustainability Research Institute; University of the Witwatersrand; Private Bag X3 WITS 2050 Johannesburg South Africa
4. British Antarctic Survey; NERC; High Cross, Madingley Road Cambridge CB3 0ET UK
5. National Antarctic Research Center, Institute of Graduate Studies; University of Malaya; 50603 Kuala Lumpur Malaysia