1. Department of Pathology and Biology of Diseases Graduate School of Medicine Kyoto University Kyoto Japan
2. Department of Hematology‐Oncology Osaka University Graduate School of Medicine Osaka Japan
3. Laboratory of Bioimaging and Cell Signaling Graduate School of Medicine Kyoto University Kyoto Japan
4. Laboratory for Mouse Genetic Engineering Quantitative Biology Center RIKEN Suita, Osaka Japan
5. Research Institute National Cerebral and Cardiovascular Center Suita, Osaka Japan