1. Department of Zoology and Entomology Rhodes University Grahamstown South Africa
2. Earth and Life Institute, Biodiversity Research Centre UCLouvain Louvain‐la‐Neuve Belgium
3. Department of Biochemistry and Microbiology Rhodes University Grahamstown South Africa
4. UMR‐CNRS 6553 EcoBio Université de Rennes 1 Rennes France