1. Faculty of Environmental & Symbiotic Sciences; Prefectural University of Kumamoto; 3-1-100 Tsukide Kumamoto 862-8502 Japan
2. Department of Marine Science and Technology; Fukui Prefectural University; 1-1 Gakuen-cho Obama Fukui 917-0003 Japan
3. National Institute for Environmental Studies; 16-2 Onogawa Tsukuba 305-8506 Japan
4. Tohoku National Fisheries Research Institute; 3-27-5 Shinhama-cho Shiogama Miyagi 985-0001 Japan