1. Institute of Marine Biology; National Taiwan Ocean University; Keelung 20224 Taiwan R.O.C.
2. Department of Marine Organism Taxonomy and Phylogeny; Institute of Oceanology; Chinese Academy of Sciences; Qingdao 266701 China
3. National Research Institute of Fisheries and Environment of Inland Sea; Fisheries Research Agency; Hatsukaichi Hiroshima 739-0452 Japan