1. Key Laboratory of Tropical Marine Bio-resources and Ecology; Guangdong Provincial Key Laboratory of Applied Marine Biology; South China Sea Institute of Oceanology; Chinese Academy of Sciences; Guangzhou China
2. South China Sea Bio-Resource Exploitation and Utilization Collaborative Innovation Center; Guangzhou China
3. Fisheries College; Ocean University of China; Qingdao China
4. Engineering Research Center of Shellfish Culture and Breeding of Liaoning Province; College of Fisheries and Life Science; Dalian Ocean University; Dalian China