1. Beijing Advanced Innovation Center for Tree Breeding by Molecular Design Beijing Forestry University No. 35 Qinghua East Road Beijing100083China
2. National Engineering Laboratory for Tree Breeding College of Biological Sciences and Technology Beijing Forestry University No. 35 Qinghua East Road Beijing100083China
3. Key Laboratory of Genetics and Breeding in Forest Trees and Ornamental Plants College of Biological Sciences and Technology Beijing Forestry University No. 35 Qinghua East Road Beijing100083China
4. Department of Plant Biology Linnean Center for Plant Biology Swedish University of Agricultural Sciences Box 7080 UppsalaSE‐750 07Sweden
5. Department of Forest and Conservation Sciences Faculty of Forestry The University of British Columbia Vancouver BCV6T 1Z4Canada