1. State Key Laboratory of Biocontrol; Institute of Aquatic Economic Animals and the Guangdong Province Key Laboratory for Aquatic Economic Animals; Sun Yat-Sen University; Guangzhou China
2. Department of life science; Hunan Normal University; Changsha China
3. Guangdong Daya Bay Fisheries Development Center; Huizhou China
4. College of Ocean; Hainan University; Haikou China