1. Department of Animal and Plant Sciences; University of Sheffield; Sheffield UK
2. Institute for Advanced Study in Toulouse; Toulouse France
3. University of Montpellier; Montpellier France
4. Institute des Sciences de l'Evolution-Montpellier; Centre National de la Recherche Scientifique; Montpellier Cedex 5 France
5. Department of Biology; University of Turku; Turku Finland