1. Ecology and Evolutionary Biology Princeton University Princeton NJ 08544 USA
2. Department of Biostatistics UCLA Fielding School of Public Health University of California, Los Angeles Los Angeles CA 90095 USA
3. Molecular, Cell and Developmental Biology University of California, Los Angeles Los Angeles CA 90095 USA
4. Department of Biological Sciences Wayne State University Detroit MI 48085 USA
5. Yale School of Management Yale University New Haven CT 06511 USA
6. Yellowstone Center for Resources National Park Service Yellowstone National Park WY 82190 USA
7. National Human Genome Research Institute National Institutes of Health Bethesda MD 20892 USA
8. Ecology and Evolutionary Biology University of California, Los Angeles Los Angeles CA 90095 USA
9. Department of Human Genetics and Biomathematics David Geffen School of Medicine at UCLA Los Angeles CA 90095 USA