1. Guangdong Provincial Key Laboratory of Applied Botany, South China Botanical Garden Chinese Academy of Sciences Guangzhou 510650 China
2. College of Forestry and Landscape Architecture South China Agricultural University Guangzhou 510642 China
3. State Key Laboratory of Systematic and Evolutionary Botany, Institute of Botany Chinese Academy of Sciences Beijing 100093 China
4. Komarov Botanical Institute, Russian Academy of Sciences Professor Popov Street 2 Saint Petersburg 197376 Russia
5. Guangdong Eco‐Engineering Polytechnic Guangzhou 510520 China
6. Department of Biology, Faculty of Science Selçuk University Konya 42031 Turkey
7. Institute of Plant and Animal Ecology, Ural Branch, Russian Academy of Sciences Yekaterinburg 620144 Russia
8. Department of Botany National Museum of Natural History, MRC 166, Smithsonian Institution Washington DC 20013‐7012 USA