1. Key Laboratory for Plant Diversity and Biogeography of East Asia; Kunming Institute of Botany; Chinese Academy of Sciences; 132 Lanhei Road Kunming 650201 China
2. University of Chinese Academy of Sciences; Beijing 100049 China
3. School of Life Science; Yunnan Normal University; Kunming 650500 China
4. College of Forestry; Southwest Forestry University; Kunming 650224 China
5. Department of Ecology & Evolution; University of Salzburg; Hellbrunnerstrasse 34 A-5020 Salzburg Austria