1. Museum of Comparative Zoology; Department of Organismic and Evolutionary Biology; Harvard University; Cambridge Massachusetts 02138 USA
2. Division of Invertebrate Zoology; American Museum of Natural History; New York New York 10024 USA
3. Sackler Institute for Comparative Genomics; American Museum of Natural History; New York New York 10024 USA
4. Institute of Biological Sciences; Science Faculty; University of Malaya; Kuala Lumpur 50603 Malaysia