1. School of Biological Sciences; The University of Queensland; Brisbane Qld 4072 Australia
2. Allan Wilson Centre for Molecular Ecology and Evolution; New Zealand Institute for Advanced Study; Institute of Natural and Mathematical Sciences; Massey University; Auckland 0745 New Zealand
3. Auckland War Memorial Museum; Tāmaki Paenga Hira; Auckland 1142 New Zealand
4. CIBIO/InBIO - Centro de Investigação em Biodiversidade e Recursos Genéticos da Universidade do Porto; R. Padre Armando Quintas 4485-661 Vairão Portugal