1. National Agricultural Research Center; National Agriculture and Food Research Organization; 3-1-1 Kannondai; Tsukuba; Ibaraki; 305-8666; Japan
2. Laboratory of Pathogen-Plant Interactions; Division of Applied Bioscience; Research Faculty of Agriculture; Hokkaido University; Kita 9, Nishi 9, Kita-ku; Sapporo; 060-8589; Japan