1. Department of Environmental Conservation University of Massachusetts Amherst Amherst Massachusetts 01003 USA
2. Department of Biology University of Massachusetts Amherst Amherst Massachusetts 01003 USA
3. Department of Applied Ecology North Carolina State University Raleigh North Carolina 27695 USA
4. Royal Botanic Gardens Kew Green, Kew, Richmond Surrey TW9 3AE UK
5. Natural Resources Institute University of Greenwich Kent ME4 4TB UK