1. Joint Research Unit Ecology of Guiana Forests; CNRS-UMR 8172 Campus agronomique, BP 316, 97379 Kourou cedex France
2. Laboratory of Evolution and Biological Diversity; Bât. 4R1, Université Paul Sabatier; 118 route de Narbonne 31062 Toulouse cedex 9 France
3. Laboratory of Functional Ecology and Environment; Bât. 4R1, Université Paul Sabatier; 118 route de Narbonne 31062 Toulouse cedex 9 France
4. Center for Functional and Evolutive Ecology; Université Montpellier 2; 1919 route de Mende 34293 Montpellier France