1. Key Laboratory of Vegetation Restoration and Management of Degraded Ecosystems Guangdong Provincial Key Laboratory of Applied Botany South China Botanical Garden Chinese Academy of Sciences Guangzhou China
2. Southern Marine Science and Engineering Guangdong Laboratory (Guangzhou) Guangzhou China
3. Forest Ecology Research Center Research Institute of Tropical Forestry Chinese Academy of Forestry Guangzhou China
4. School of Biological Sciences University of Nebraska Lincoln NE USA
5. Center for Plant Science Innovation University of Nebraska Lincoln NE USA