1. Centre for Ecology and Conservation College of Life & Environmental Sciences University of Exeter Penryn Campus Cornwall TR10 9EZ UK
2. Department of Entomology North Carolina State University Gardner Hall Raleigh North Carolina 27695‐7613 USA
3. Laboratory of Evolutionary Ecology Graduate School of Environmental Science Okayama University Tsushima‐naka 111 Okayama 700‐8530 Japan