1. Xishuangbanna Tropical Botanical Garden Chinese Academy of Sciences Centre for Integrated Conservation, Community Ecology and Conservation Menglun 666303 Yunnan China
2. CIRAD UMR AMAP TA 40/PS2 34398 Montpellier Cedex 5 France
3. Department of Zoology Nelson Mandela Metropolitan University Port Elizabeth Province of Eastern Cape South Africa
4. Centre d'Ecologie Fonctionnelle et Evolutive UMR 5175 Centre National de la Recherche Scientifique (CNRS) Université de Montpellier Université Paul Valéry Montpellier Ecole Pratiques des Hautes Etudes (EPHE) 1919 Route de Mende 34293 Montpellier Cedex 5 France