1. Department of Biological Sciences; Northern Arizona University; Flagstaff Arizona 86011 USA
2. Tree Improvement Branch; British Columbia Ministry of Forests, Lands and Natural Resource Operations; Kalamalka Forestry Centre; 3401 Reservoir Road Vernon British Columbia V1B 2C7 Canada
3. Merriam-Powell Center for Environmental Research; Northern Arizona University; Flagstaff Arizona 86011 USA
4. School of Forestry; Northern Arizona University; Flagstaff Arizona 86011 USA