1. Institut des Sciences de l'Evolution Université de Montpellier CNRS, IRD, EPHE CC 065 Place Eugène Bataillon 34095 Montpellier Cedex 05 France
2. Resource Ecology Group Wageningen University P.O. Box 47 NL‐6700 AA Wageningen the Netherlands
3. Aquatic Ecology and Water Quality Management Group Wageningen University P.O. Box 47 NL‐6700 AA Wageningen the Netherlands