1. Biological and Environmental Sciences University of Stirling Stirling FK9 4LA UK
2. Centre of Ecology and Conservation College of Life and Environmental Sciences University of Exeter Penryn Campus Penryn, CornwallTR10 9FE UK
3. Department of Respiratory, Inflammation and Autoimmunity MedImmune Ltd. Abington CB21 6GH UK
4. School of Science and Health Western Sydney University Locked Bag 1797 Penrith New South Wales2751 Australia