1. University Bordeaux 1; UMR CNRS 5805 EPOC; Avenue des Facultés; 33405 Talence France
2. The James Hutton Institute; Craigiebuckler Aberdeen AB15 8QH UK
3. Department of Biology; York University; 4700 Keele Street Toronto M3J 1P3 Ontario Canada
4. Division of Biological Sciences; University of Montana; Missoula Montana 59812 USA